The common ostrich (Struthio camelus) or simply ostrich, is a species of large flightless bird native to certain large areas of Africa. It is one of two extant species of ostriches, the only living members of the genus Struthio in the ratite order of birds. The other is the Somali ostrich (Struthio molybdophanes), which was recognized as a distinct species by BirdLife International in 2014 having been previously considered a very distinctive subspecies of ostrich.
The common ostrich belongs to the order Struthioniformes. Struthioniformes contained all the ratites, such as the kiwis, emus, rheas, and cassowaries. However, recent genetic analysis has found that the group is not monophyletic, as it is paraphyletic with respect to the tinamous, so the ostriches are classified as the only members of the order. Phylogenetic studies have shown that it is the sister group to all other members of Palaeognathae and thus the flighted tinamous are the sister group to the extinct moa. It is distinctive in its appearance, with a long neck and legs, and can run for a long time at a speed of 55 km/h (34 mph) with short bursts up to about 70 km/h (43 mph), the fastest land speed of any bird.The common ostrich is the largest living species of bird and lays the largest eggs of any living bird (the extinct elephant birds of Madagascar and the giant moa of New Zealand laid larger eggs).
The common ostrich's diet consists mainly of plant matter, though it also eats invertebrates. It lives in nomadic groups of 5 to 50 birds. When threatened, the ostrich will either hide itself by lying flat against the ground, or run away. If cornered, it can attack with a kick of its powerful legs. Mating patterns differ by geographical region, but territorial males fight for a harem of two to seven females.
The common ostrich is farmed around the world, particularly for its feathers, which are decorative and are also used as feather dusters. Its skin is used for leather products and its meat is marketed commercially, with its leanness a common marketing point.
Description & appearance
Common ostriches usually weigh from 63 to 145 kilograms (139–320 lb), or as much as two adult humans. The Masai ostriches of East Africa (S. c. massaicus) averaged 115 kg (254 lb) in males and 100 kg (220 lb) in females, while the nominate subspecies, the North African ostrich (S. c. camelus), was found to average 111 kg (245 lb) in unsexed adults. Exceptional male ostriches (in the nominate subspecies) can weigh up to 156.8 kg (346 lb). At sexual maturity (two to four years), male common ostriches can be from 2.1 to 2.8 m (6 ft 11 in to 9 ft 2 in) in height, while female common ostriches range from 1.7 to 2.0 m (5 ft 7 in to 6 ft 7 in) tall. New chicks are fawn in color, with dark brown spots. During the first year of life, chicks grow at about 25 cm (9.8 in) per month. At one year of age, common ostriches weigh approximately 45 kilograms (99 lb). Their lifespan is up to 40–45 years.
The feathers of adult males are mostly black, with white primaries and a white tail. However, the tail of one subspecies is buff. Females and young males are grayish-brown and white. The head and neck of both male and female ostriches is nearly bare, with a thin layer of down. The skin of the female's neck and thighs is pinkish gray, while the male's is gray or pink dependent on subspecies.
The long neck and legs keep their head up to 2.8 m (9 ft) above the ground, and their eyes are said to be the largest of any land vertebrate: 50 mm (2.0 in) in diameter; helping them to see predators at a great distance. The eyes are shaded from sunlight from above. However, the head and bill are relatively small for the birds' huge size, with the bill measuring 12 to 14.3 cm (4.7 to 5.6 in).
Their skin varies in color depending on the subspecies, with some having light or dark gray skin and others having pinkish or even reddish skin. The strong legs of the common ostrich are unfeathered and show bare skin, with the tarsus (the lowest upright part of the leg) being covered in scales: red in the male, black in the female. The tarsus of the common ostrich is the largest of any living bird, measuring 39 to 53 cm (15 to 21 in) in length. The bird has just two toes on each foot (most birds have four), with the nail on the larger, inner toe resembling a hoof. The outer toe has no nail. The reduced number of toes is an adaptation that appears to aid in running, useful for getting away from predators. Common ostriches can run at a speed over 70 km/h (43 mph) and can cover 3 to 5 m (9.8 to 16.4 ft) in a single stride. The wings reach a span of about 2 metres (6 ft 7 in), and the wing chord measurement of 90 cm (35 in) is around the same size as for the largest flying birds.
The feathers lack the tiny hooks that lock together the smooth external feathers of flying birds, and so are soft and fluffy and serve as insulation. Common ostriches can tolerate a wide range of temperatures. In much of their habitat, temperatures vary as much as 40 °C (72 °F) between night and day. Their temperature control relies in part on behavioral thermoregulation. For example, they use their wings to cover the naked skin of the upper legs and flanks to conserve heat, or leave these areas bare to release heat. The wings also function as stabilizers to give better maneuverability when running. Tests have shown that the wings are actively involved in rapid braking, turning and zigzag maneuvers. They have 50–60 tail feathers, and their wings have 16 primary, four alular and 20–23 secondary feathers.
The common ostrich's sternum is flat, lacking the keel to which wing muscles attach in flying birds. The beak is flat and broad, with a rounded tip. Like all ratites, the ostrich has no crop, and it also lacks a gallbladder. They have three stomachs, and the caecum is 71 cm (28 in) long. Unlike all other living birds, the common ostrich secretes urine separately from faeces. All other birds store the urine and faeces combined in the coprodeum, but the ostrich stores the faeces in the terminal rectum. They also have unique pubic bones that are fused to hold their gut. Unlike most birds, the males have a copulatory organ, which is retractable and 20 cm (8 in) long. Their palate differs from other ratites in that the sphenoid and palatal bones are unconnected.
Common ostriches normally spend the winter months in pairs or alone. Only 16 percent of common ostrich sightings were of more than two birds.[9] During breeding season and sometimes during extreme rainless periods ostriches live in nomadic groups of five to 100 birds (led by a top hen) that often travel together with other grazing animals, such as zebras or antelopes. Ostriches are diurnal, but may be active on moonlit nights. They are most active early and late in the day. The male common ostrich territory is between 2 and 20 km2 (0.77 and 7.72 sq mi).
With their acute eyesight and hearing, common ostriches can sense predators such as lions from far away. When being pursued by a predator, they have been known to reach speeds in excess of 70 km/h (43 mph), and can maintain a steady speed of 50 km/h (31 mph), which makes the common ostrich the world's fastest two-legged animal. When lying down and hiding from predators, the birds lay their heads and necks flat on the ground, making them appear like a mound of earth from a distance, aided by the heat haze in their hot, dry habitat.
When threatened, common ostriches run away, but they can cause serious injury and death with kicks from their powerful legs.Their legs can only kick forward. Distribution & habitat
Common ostriches formerly occupied Africa north and south of the Sahara, East Africa,
Africa South of the rainforest belt, and much of Asia Minor.Today common ostriches prefer open land and are native to the savannas and Sahel of Africa, both north and south of the equatorial forest zone. In southwest Africa they inhabit the semi-desert or true desert. Farmed common ostriches in Australia have established feral populations. The Arabian ostriches in the Near and Middle East were hunted to extinction by the middle of the 20th century. Attempts to reintroduce the common ostrich into Israel have failed. Common ostriches have occasionally been seen inhabiting islands on the Dahlak Archipelago, in the Red Sea near Eritrea.
Research conducted by the Birbal Sahni Institute of Palaeobotany in India found molecular evidence that ostriches lived in India 25,000 years ago. DNA tests on fossilized eggshells recovered from eight archaeological sites in the states of Rajasthan, Gujarat and Madhya Pradesh found 92% genetic similarity between the eggshells and the North African ostrich, so these could have been fairly distant relatives.
Food
They mainly feed on seeds, shrubs, grass, fruit and flowers; occasionally they also eat insects such as locusts. Lacking teeth, they swallow pebbles that act as gastroliths to grind food in the gizzard. When eating, they will fill their gullet with food, which is in turn passed down their esophagus in the form of a ball called a bolus. The bolus may be as much as 210 ml (7.1 US fl oz). After passing through the neck (there is no crop) the food enters the gizzard and is worked on by the aforementioned pebbles. The gizzard can hold as much as 1,300 g (46 oz), of which up to 45% may be sand and pebbles. Common ostriches can go without drinking for several days, using metabolic water and moisture in ingested plants, but they enjoy liquid water and frequently take baths where it is available. They can survive losing up to 25% of their body weight through dehydration. Breeding & mating
Common ostriches become sexually mature when they are 2 to 4 years old; females mature about six months earlier than males. As with other birds, an individual may reproduce several times over its lifetime. The mating season begins in March or April and ends sometime before September. The mating process differs in different geographical regions. Territorial males typically boom in defense of their territory and harem of two to seven hens; the successful male may then mate with several females in the area, but will only form a pair bond with a 'major' female.
The cock performs with his wings, alternating wing beats, until he attracts a mate. They will go to the mating area and he will maintain privacy by driving away all intruders. They graze until their behavior is synchronized, then the feeding becomes secondary and the process takes on a ritualistic appearance. The cock will then excitedly flap alternate wings again, and start poking on the ground with his bill. He will then violently flap his wings to symbolically clear out a nest in the soil. Then, while the hen runs a circle around him with lowered wings, he will wind his head in a spiral motion. She will drop to the ground and he will mount for copulation. Common ostriches raised entirely by humans may direct their courtship behavior not at other ostriches, but toward their human keepers. Chicks, juveniles & raise
The female common ostrich lays her fertilized eggs in a single communal nest, a simple pit, 30 to 60 cm (12–24 in) deep and 3 m (9.8 ft) wide, scraped in the ground by the male. The dominant female lays her eggs first, and when it is time to cover them for incubation she discards extra eggs from the weaker females, leaving about 20 in most cases. A female common ostrich can distinguish her own eggs from the others in a communal nest.[56] Ostrich eggs are the largest of all eggs, though they are actually the smallest eggs relative to the size of the adult bird — on average they are 15 cm (5.9 in) long, 13 cm (5.1 in) wide, and weigh 1.4 kilograms (3.1 lb), over 20 times the weight of a chicken's egg and only 1 to 4% the size of the female. They are glossy cream-colored, with thick shells marked by small pits.
The eggs are incubated by the females by day and by the males by night. This uses the coloration of the two sexes to escape detection of the nest, as the drab female blends in with the sand, while the black male is nearly undetectable in the night. The incubation period is 35 to 45 days, which is rather short compared to other ratites. This is believed to be the case due to the high rate of predation. Typically, the male defends the hatchlings and teaches them to feed, although males and females cooperate in rearing chicks. Fewer than 10% of nests survive the 9 week period of laying and incubation, and of the surviving chicks, only 15% of those survive to 1 year of age. However, among those common ostriches who survive to adulthood, the species is one of the longest-living bird species. Common ostriches in captivity have lived to 62 years and 7 months.